gs \
  -o OUTPUT.pdf \
  -sDEVICE=pdfwrite \
  -dDownsampleColorImages=true \
  -dDownsampleGrayImages=true \
  -dDownsampleMonoImages=true \
  -dColorImageResolution=72 \
  -dGrayImageResolution=72 \
  -dMonoImageResolution=72 \
  -dColorImageDownsampleThreshold=1.0 \
  -dGrayImageDownsampleThreshold=1.0 \
  -dMonoImageDownsampleThreshold=1.0 \
   YOUR_INPUT_PDF_FILE.pdf